Darwin also recognised that variations differ in degree, even as they do
in kind. He perceived that some variations are much more pronounced than
others. He recognised the distinction between what are now known as
continuous and discontinuous variations. The former are slight departures
from the normal; the latter are considerable deviations from the mean or
mode; great jumps, as it were, taken by nature, as, for example, the pea
and the rose combs of fowls, which were derived from the normal single
comb.
Monstrosities
“At long intervals of time,” wrote Darwin, “out of millions of
individuals reared in the same country and fed on nearly the same food,
deviations of structure so strongly pronounced as to deserve to be called
monstrosities arise, but monstrosities cannot be separated by any
distinct line from slighter variations.” Therefore it is evident that he
regarded the difference between continuous and discontinuous variations
as not one of kind, but merely of degree. To the discontinuous variations
Darwin attached very little importance from an evolutionary point of
view. He looked upon them as something abnormal.
“It may be doubted,” he wrote, “whether such sudden and considerable
deviations of structure such as we occasionally see in our domestic
productions, more especially with plants, are ever permanently propagated
in a state of nature. Almost every part of every organic being is so
beautifully related to its complex conditions of life that it seems as
improbable that any part should have been suddenly produced perfect, as
that a complex machine should have been invented by a man in a perfect
state. Under domestication monstrosities sometimes occur which resemble
normal structures in widely different animals. Thus pigs have
occasionally been born with a sort of proboscis, and if any wild species
of the same genus had naturally possessed a proboscis, it might have been
argued that this had appeared as a monstrosity; but I have as yet failed
to find, after diligent search, cases of monstrosities resembling normal
structures in nearly allied forms, and these alone bear on the question.
If monstrous forms of this kind ever do appear in a state of nature and
are capable of reproduction (which is not always the case), as they occur
rarely and singly, their preservation would depend on unusually
favourable circumstances. They would, also, during the first and
succeeding generations cross with the ordinary form, and thus their
abnormal character would almost inevitably be lost.” But, in a later
edition of the _Origin of Species_, Darwin seems to contradict the above
assertion: “It should not, however, be overlooked that certain rather
strongly marked variations, which no one would rank as mere individual
differences, frequently recur owing to a similar organisation being
similarly acted on—of which fact numerous instances could be given with
our domestic productions.
Public-domain text, read in full here on John Shaqi.
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