In such cases, if the varying individual did
not actually transmit to its offspring its newly acquired character, it
would undoubtedly transmit to them, as long as the existing conditions
remained the same, a still stronger tendency to vary in the same manner.
There can also be little doubt that the tendency to vary in the same
manner has often been so strong that all the individuals of the same
species have been similarly modified without the aid of any form of
selection. Or only a third, fifth, or tenth part of the individuals may
have been thus affected, of which fact several instances could be given.
Thus Graba estimates that about one-fifth of the guillemots in the Faroe
islands consist of a variety so well marked, that it was formerly ranked
as a distinct species under the name _Uria lacrymans_. In cases of this
kind, if the variation were of a beneficial nature, the original form
would soon be supplanted by the modified form, through the survival of
the fittest.” Here we seem to have a plain statement of the origin of new
forms by mutation.
Minute Variations
Again, we read (page 34): “Some variations useful to him (_i.e._ man)
have probably arisen suddenly, or by one step; many botanists, for
instance, believe that the fuller’s teasel, with its hooks, which cannot
be rivalled by any mechanical contrivance, is only a variety of the wild
Dipsacus; and this amount of change may have suddenly arisen in a
seedling. This is known to be the case with the turnspit dog.”[2] But, as
we have already said, Darwin at no time attached much importance to these
jumps made by nature as a factor in evolution. He pinned his faith to the
minute, indefinite variations which he believed could be piled up, one
upon another, so that, if allowed sufficient time, either nature or the
human breeder could, by a continued selection of these minute variations,
call into being any kind of organism. The importance of selection, he
writes, “consists in the great effect produced by the accumulation in one
direction, during successive generations, of differences absolutely
inappreciable by an uneducated eye” (page 36). On page 132 he writes: “I
can see no limit to the amount of change, to the beauty and complexity of
the coadaptations between all organic beings . . . which may have been
effected[3] in the long course of time by nature’s power of selection.”
He expressly states, on page 149, that he sees no reason to limit the
process to the formation of genera alone.
Public-domain text, read in full here on John Shaqi.
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