Both Wasps accumulate caterpillars in their coffers, whether domes or
jars. Let us give an abstract of the bill of fare. These documents, for
all their dryness, possess a value: they will enable whoso cares to
interest himself in the Eumenes to perceive to what extent instinct
modifies the diet, according to place and season. The food is plentiful
but lacks variety. It consists of tiny caterpillars, by which I mean
the grubs of small Butterflies or Moths. This is proclaimed by the
structure, for we observe the usual caterpillar organism in the prey
selected by either Wasp. The body is composed of twelve segments, not
including the head. The first three have true legs, the next two are
legless, then come four segments with prolegs, two legless segments
and, lastly, a terminal segment with prolegs. It is exactly the same
organization which we saw in the Ammophila’s Grey Worm.
My old notes give the following description of the caterpillars found
in the nest of E. Amadei: a pale-green or, less often, yellowish body
covered with short white hairs; head wider than the front segment,
dead-black and also bristling with hairs. Length: 16 to 18 millimetres;
[6] width: about 3 millimetres. [7] It is more than a quarter of a
century since I jotted down this descriptive sketch; and today, at
Sérignan, I find in the Eumenes’ larder the same sort of game that I
saw long ago at Carpentras. Time and distance have not altered the
nature of the provisions.
I know one exception and one alone in this fidelity to the ancestral
diet. My observations mention a single dish that differs greatly from
those which accompany it. This is a caterpillar of the Looper group [8]
with only three pairs of prolegs, placed under the eighth, ninth and
twelfth segments. The body tapers slightly at either end, is contracted
at the junction of the different rings and is pale green with faint
black veinings, visible under the magnifying-glass, and a few sparse
black cilia. Length: 15 millimetres; [9] width: 2½ millimetres. [10]
E. pomiformis also has her preferences. Her game consists of small
caterpillars about 7 millimetres long by 1⅓ wide. [11] The body is pale
green, pretty sharply contracted at the junction of the segments. The
head is narrower than the rest of the body and is spotted with brown.
Pale ocellated circles are distributed in two transversal rows over the
middle segments and have a black dot in the centre, surmounted by a
black cilium. On the third and fourth and also on the penultimate
segment, each circle has two black dots and two cilia. This is the
rule.
The exception is supplied by two head of game in the whole course of my
observations. These two had a pale yellow body, with five longitudinal
brick-red stripes and a few very rare cilia. Head and prothorax brown
and shiny; length and diameter as above.
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