The North American Slime-Moulds: A Descriptive List of All Species of Myxomycetes Hitherto Reported from the Continent of North America, with Notes on Some Extra-Limital Species — John Shaqi
The North American Slime-Moulds: A Descriptive List of All Species of Myxomycetes Hitherto Reported from the Continent of North America, with Notes on Some Extra-Limital SpeciesMacbride, Thomas H. (Thomas Huston)
Science
The North American Slime-Moulds: A Descriptive List of All Species of Myxomycetes Hitherto Reported from the Continent of North America, with Notes on Some Extra-Limital Species
Macbride, Thomas H. (Thomas Huston)
Myxomycetes
Under certain conditions, low temperature, lack of moisture, the
plasmodium may pass into a resting phase, when it masses itself in heaps
and may become quite dry in lumps of considerable size, and so await the
return of favorable conditions when former activity is quickly resumed.
Sometimes the larger plasmodia pass into the resting phase by undergoing
a very peculiar change of structure. In ordinary circumstances the
abundant free nuclei demonstrable in the plasmodium afford the only
evidence of cellular organization. In passing now into the condition of
rest, the whole protoplasmic mass separates simultaneously into numerous
definite polyhedral or parenchymatous cells, each with a well-developed
_cellulose wall_.[4] When the conditions essential to activity are
restored, the walls disappear, the cellulose is resorbed, and the
plasmodium resumes its usual habit and structure.
The plasmodial phase of the slime-mould, like the hyphal phase of the
fungus, may continue a long time; for months, possibly for years. The
reason for making the latter statement will presently appear. But
however long or short the plasmodial phase continue, the time of fruit,
the reproductive phase, at length arrives. When this time comes, induced
partly by a certain maturity in the organism itself, partly no doubt by
the trend of external conditions, the plasmodium no longer as before
evades the light, but pushes to the surface, and appears usually in some
elevated or exposed position, the upper side of the log, the top of the
stump, the upper surface of its habitat, whatever that may be; or even
leaves its nutrient base entirely and finds lodging on some neighboring
object. In such emergency the stems and leaves of flowering plants are
often made to serve, and even fruits and flowers afford convenient
resting places. The object now to be attained is not the formation of
fruit alone, but likewise its speedy desiccation and the prompt
dispersal of the perfected spores. Nothing can be more interesting than
to watch the slime-mould as its plasmodium accomplishes this its last
migration. If hitherto its habitat has been the soft interior of a
rotten log, it now begins to ooze out in all directions, to well up
through the crevices of the bark as if pushed by some energy acting in
the rear, to stream down upon the ground, to flow in a hundred tiny
streams over all the region round about, to climb all stems, ascend all
branches, to the height of many inches, all to pass suddenly as if by
magic charm into one widespread, dusty field of flying spores. Or, to be
more exact, whatever the position ultimately assumed, the plasmodium
soon becomes quiescent, takes on definite and ultimate shape, which
varies greatly, almost for each species. Thus it may simply form a flat,
cake-like mass, _aethalium_, internally divided into an indefinite
number of ill-defined spore cases, sporangia; or the plasmodium may take
the form of a simple net, _plasmodiocarp_, whose cords stand out like
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