The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
sees in Pterodactyle an indication of the course that nature took in
changing the reptilian organization to that of birds and mammals. The
less important organs, those of motion, assimilate partly to those
of the bird and partly to those of the bats, but always preserve
the fundament reptile type and reptile number of bones. The skull,
fluctuating in character between the monitor and crocodile, hides its
reptile nature under the outer form of the bird, but retains the teeth.
To change the skull into a bird's skull it would only be necessary that
a few separate elements should be blended together, and that a few
peculiar bones should be removed. The length of the neck, varying only
in a few species, is a deviation from the reptile type, and indicates
an approximation to the structure of 'the bird; but the number of
the vertebræ remains constant notwithstanding the increased length.
The fundamental plan of the crocodile may be recognised in all the
important parts of the vertebræ. The body of the reptile, to be enabled
to fly, would need a larger breast and a stronger structure of the
fore-limbs. The shoulder-blade of the reptile, with its extremities
forming the glenoid cavity, is necessarily smaller and prolonged
backward, and altered to resemble that of a bird. The scapula only
formed the back part of the glenoid cavity, but it is thick and strong,
suggesting an affinity with the bats.
The breast-bone, in the form of a shield, is changing into that of a
bird; as are the ribs, which are attached in a peculiar way to the
vertebral column. It is really the strong sternum of the Chameleon,
with moveable dorsal vertebræ. The whole chest is supported by the
peculiar continuation of the wings of the pubic bones (Schambein).
The ischiac and pubic bones resemble those of the Chameleon, but the
ilium runs a little down, like that of a bird, and is only slightly
connected with two sacral vertebræ, as in reptiles, prolonging itself
a little upward and forward, as in mammals. The wings of the pubic
bones exist in the Turtle and Monitor, but of small extent; they are
also represented in the mammals by the upward development of the pubic
bones in those families, genera, and species, in which nature has
indicated by variety of shape, or peculiarities of development, or by
affinities with reptiles, quite a new type and capacity for variation
within certain limits, which is especially the case with certain
Rodents and Opossums, and Monotremes. It would not be astonishing to
find in Pterodactyles the marsupial bones. And indeed the Pterodactylus
crassirostris has a small tongue-shaped bone, probably belonging to the
pelvis. The less important part of the skeleton, the tail, is formed
precisely as in mammals, and is identical with that of the bats. Both
the thigh and shin are mammalian, and only the foot retains the same
number of parts as in reptiles.
This animal was enabled by means of the pelvic bones and the long
hind-legs to sit like the squirrels.
Public-domain text, read in full here on John Shaqi.
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