the integument better than can the general protoplasm; these are the
nerves. Other parts are modified so that they can contract or secrete
the more easily; these are the muscles and glands. The conception
of a reflex action, as it is usually stated in books on physiology,
therefore includes this idea of the differentiation of the tissues, but
all the processes that are included in the typical reflex are processes
which can be carried on by undifferentiated protoplasm.
It is also a schematic description that assumes a simplicity that does
not really exist. As a rule a reflex is initiated by the stimulation
of more than one receptor organ, and the impulses initiated may thus
reach the central nervous system by more than one path. There is no
simple shunting of the afferent impulse from the cell in which it
terminates into another nerve, when it becomes an efferent impulse;
but, instead of this, the impulse may “zigzag” through a maze of paths
in the brain or spinal cord connecting together afferent and efferent
nerves and ganglia. Further, the final part of the reflex, the muscular
contraction, is far from being a simple thing, for usually a series
of muscles are stimulated to contract, each of them at the right time
and with the right amount of force, and every contraction of a muscle
is accompanied by the relaxation of the antagonistic muscle. There are
muscles which open the eyelids and others which close them, and the
cerebral impulse which causes the levators to contract at the same time
causes the depressors to relax.
It is quite necessary to remember that the simple reflex is really
a process of much complexity and may involve many other parts and
structures than those to which we immediately direct our attention. But
leaving aside these qualifications we may usefully consider the general
characters of the reflex, regarding it as a common, automatically
performed, restricted bodily action, involving receptor organ, central
nervous organ, and effector organ. There are certain kinds of external
stimuli that continually affect our organs of sense, and there are
certain kinds of muscular and glandular activity that occur “as a
matter of course,” when these stimuli fall on our organs of sense.
The emanation from onions or the vapour of ammonia causes our eyes to
water; the smell of savoury food causes a flow of saliva; and anything
that approaches the face very rapidly causes us to close the eyes.
Reflexes are, in a way, commonly occurring, purposeful and useful
actions, and their object is the maintenance of a normal condition of
bodily functioning.
Public-domain text, read in full here on John Shaqi.
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