The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Where, as in types like these, the morphological units show an inherent
tendency to arrange themselves in a manner which is so far constant
as to give characteristic proportions, we may say that there is a
recognizable compound individuality. Considering the Thallophytes which
grow in this way apart from their kinships, and wholly with reference
to their morphological composition, we might not inaptly describe them
as pseudo-foliar.
§ 184. Another mode in which aggregation is so carried on as to produce
a compound individuality of considerable definiteness, is variously
displayed among other families of _Algæ_. When the cells, instead
of multiplying longitudinally alone, and instead of all multiplying
laterally as well as longitudinally, multiply laterally only at
particular places, they produce branched structures.
Indications of this mode of aggregation occur among the _Confervoideæ_,
as shown in Figs. 22, 23. Though, in some of the more-developed
_Algæ_ which exhibit the ramified arrangement in a higher degree, the
component cells are, like those of the lower _Algæ_, united together
end to end, in such way as but little to obscure their separate
forms, as in _Cladophora Hutchinsiæ_, Fig. 31; they nevertheless
evince greater subordination to the whole of which they are parts,
by arranging themselves more methodically. Still further pronounced
becomes the compound individuality when, while the component cells of
the branches unite completely into jointed cylinders, the component
cells of the stem form an axis distinguished by its relative thickness
and complexity. Such types of structures are indicated by Figs. 32,
33--figures representing small portions of plants which are quite
tree-like in their entire outlines. On examining Figs. 34, 35, 36,
which show the structures of the stems in these types, it will be
seen, too, that the component cells in becoming more coherent,
have undergone changes of form which obscure their individualities
more than before. Not only are they much elongated, but they are so
compressed as to be prismatic rather than cylindrical. This structure,
besides displaying integration of the morphological units carried on
in two directions instead of one; and besides displaying this higher
integration in the greater merging of the individualities of the
morphological units in the general individuality; also displays it in
the more pronounced subordination of the branches and branchlets to the
main stem. This differentiation and consolidation of the stem, brings
all the secondary growths into more marked dependence; and so renders
the individuality of the aggregate more decided.
[Illustration: Figs. 31–36.]
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