The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 183. An approach towards plants which unite these characters, may be
traced in such forms as _Bangia ciliaris_, Fig. 24. The multiplication
of cells here takes place, not in a longitudinal direction only, but
also in a transverse direction; and the transverse multiplication being
greater towards the middle of the frond, there results a difference
between the middle and the two extremities--a character which, in
a feeble way, unites all the parts into a whole. Even this slight
individuation is, however, very indefinitely marked; since, as shown by
the figures, the lateral multiplication of cells does not go on in a
precise manner.
[Illustration: Fig. 24.]
From some such type as this there appear to arise, through slight
differences in the modes of growth, two closely-allied groups of
plants, having individualities somewhat more pronounced. If, while
the cells multiply longitudinally, their lateral multiplication goes
on in one direction only, there results a flat surface, as in the
genus _Ulva_ (Sea-lettuce) or in the upper part of the thallus of
_Enteromorpha Linza_, Fig. 25; or where the lateral multiplication is
less uniform in its rate, in types like Fig. 26. But where the lateral
multiplication occurs in two directions transverse to one another, a
hollow frond may be produced--sometimes irregularly spheroidal, and
sometimes irregularly tubular; as in _Enteromorpha intestinalis_,
Fig. 27. And often, as in _Enteromorpha compressa_, Fig. 28, and
other species, this tubular frond becomes branched. Figs. 29 and 30
are magnified portions of such fronds, showing the simple cellular
aggregation which allies them with the preceding forms.
[Illustration: Figs. 25–30.]
In the common _Fuci_ of our coasts, other and somewhat higher stages
of this integration are displayed. We have fronds preserving something
like constant breadths and dividing dichotomously with approximate
regularity. Though the subdivisions so produced are not to be regarded
as separate fronds, but only as extensions of one frond, they
foreshadow a higher degree of composition; and by the comparatively
methodic way in which they are united, give to the aggregate a more
definite, as well as a more complex, individuality. Many of the higher
lichens exhibit an analogous advance. While in the lowest lichens, the
different parts of the thallus are held together only by being all
attached to the supporting surface, in the higher lichens the thallus
is so far integrated that it can support itself by attachment to such
surface at one point only. And then, in still more developed kinds,
we find the thallus assuming a dichotomously-branched form, and so
gaining a more specific character as well as greater size.
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