The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
But among certain of the highest _Algæ_, we _do_ find something more
than this union of the pseud-axial with the pseudo-foliar structure.
In addition to pseud-axes of comparative complexity; and in addition
to pseudo-folia that are like leaves, not only in their general shapes
but in having mid-ribs and even veins; there are the beginnings of
a higher stage of integration. Figs. 38, 39, and 40, show some of
the steps. In _Rhodymenia palmata_, Fig. 38, the parent-frond is
comparatively irregular in form, and without a mid-rib; and along with
this very imperfect integration, we see that the secondary fronds
growing from the edges are distributed very much at random, and are by
no means specific in their shapes. A considerable advance is displayed
by _Phyllophora rubens_, Fig. 39. Here the frond, primary, secondary,
or tertiary, betrays some approach towards regularity in both form
and size; by which, as also by its partially-developed mid-rib, there
is established a more marked individuality; and at the same time,
the growth of the secondary fronds no longer occurs anywhere on the
edge, in the same plane as the parent-frond, but from the surface at
specific places. _Delesseria sanguinea_, Fig. 40, illustrates a much
more definite arrangement of the same kind. The fronds of this plant,
quite regularly shaped, have their parts decidedly subordinated to the
whole; and from their mid-ribs grow other fronds which are just like
them. Each of these fronds is an organized group of those morphological
units which we distinguish as aggregates of the first order. And in
this case, two or more such aggregates of the second order, well
individuated by their forms and structures, are united together; and
the plant composed of them is thus rendered, in so far, an aggregate of
the third order.
Just noting that in certain of the most developed _Algæ_, as the
_Sargassum_, or common gulf-weed, this tertiary degree of composition
is far more completely displayed, so as to produce among Thallophytes a
type of structure closely simulating that of the higher plants, let us
now pass to the consideration of these higher plants.
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