The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 186. Having the surface of the soil for a support and the air for a
medium, terrestrial plants are mechanically circumstanced in a manner
widely different from that in which aquatic plants are circumstanced.
Instead of being buoyed up by a surrounding fluid of specific gravity
equal to their own, they have to erect themselves into a rare fluid
which yields no appreciable support. Further, they are dissimilarly
conditioned in having two sources of nutriment in place of one. Unlike
the _Algæ_, which derive all the materials for their tissues from
the water bathing their entire surfaces, and use their roots only
for attachment, most of the plants which cover the Earth’s surface,
absorb part of their food through their imbedded roots and part through
their exposed leaves. These two marked unlikenesses in the relations
to surrounding conditions, profoundly affect the respective modes of
growth. We must duly bear them in mind while studying the further
advance of composition.
[Illustration: Figs. 41–44.]
[Illustration: Figs. 45–49.]
The class of plants to which we now turn--that of the
_Archegoniatæ_--is nearly related by its lower members to the classes
above dealt with: so much so, that some of the inferior liverworts are
quite licheniform, and are often mistaken for lichens. Passing over
these, let us recommence our synthesis with such members of the class
as repeat those indications of progress towards a higher composition,
which we have just observed among the more-developed _Algæ_. The
_Jungermanniaceæ_ furnish us with a series of types, clearly indicating
the transition from an aggregate of the second order to an aggregate
of the third order. Figs. 41 and 42, indicate the structure among the
lowest of this group. Here there is but an incomplete development of
the second order of aggregate. The frond grows as irregularly as the
thallus of a lichen: it is indefinite in size and outline, spreading
hither or thither as the conditions favour. Moreover, it lacks the
differentiations required to subordinate its parts to the whole: it
is uniformly cellular, having neither mid-rib nor veins; and it puts
out rootlets indifferently from all parts of its under surface. In
Fig. 43, _Pellia epiphylla_, we have an advance on this type. There
is here, as shown in the transverse section, Fig. 44, a thickening
of the frond along its central portion, producing something like an
approach towards a mid-rib; and from this the rootlets are chiefly
given off. The outline, too, is much less irregular; whence results
greater distinctness of the individuality. A further step is displayed
in _Metzgeria furcata_, Fig. 45. The frond of this plant, comparatively
well integrated by the distribution of its substance around a decided
mid-rib, and by its comparatively-definite outlines, produces secondary
fronds. There is what is called proliferous growth; and occasionally,
as shown in Fig. 46, representing an enlarged portion, the growth is
doubly-proliferous.
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