The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
but the component leaflets of the one, are usually much larger than
those of the other. The like contrasts are still more marked between
triple leaves and simple leaves. This connection of decreasing size
with decreasing composition, is conspicuous in the series of figures:
the differences shown being not nearly so great as may be frequently
observed. Confirmation may be drawn from the fact that when the leading
shoot is broken or arrested in its growth, the shoots it gives off
(provided they are given off after the injury), and into which its
checked currents of sap are thrown, produce leaves of five leaflets
where ordinarily leaves of three leaflets occur. Of course incidental
circumstances, as variations in the amounts of sunshine, or of rain,
or of matter supplied to the roots, are ever producing changes in the
state of the plant as a whole; and by thus affecting the nutrition of
its leaf-buds at the times of their formation, cause irregularities
in the relations of size and composition above described. But taking
these causes into account, it is abundantly manifest that a leaf-bud of
the bramble will develop into a simple leaf or into a leaf compounded
in different degrees, according to the quantity of assimilable matter
brought to it at the time when the rudiments of its structure are
being fixed. And on studying the habits of other plants--on observing
how annuals that have compound leaves usually bear simple leaves at
the outset, when the assimilating surface is but small; and how, when
compound-leaved plants in full growth bear simple leaves in the midst
of compound ones, the relative smallness of such simple leaves shows
that the buds from which they arose were ill-supplied with sap; it
will cease to be doubted that a foliar organ may be metamorphosed into
a group of foliar organs, if furnished, at the right time, with a
quantity of matter greater than can be readily organized round a single
centre of growth. An examination of the transitions through which
a compound leaf passes into a doubly-compound leaf, as seen in the
various intermediate forms of leaflets in Fig. 65, will further enforce
this conclusion.
[Illustration: Figs. 50–64.]
[Illustration: Fig. 65.]
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