The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Must we then conclude that as cells, or morphological units, are
integrated into a unit of a higher order, which we call a thallus
or frond; so, by the integration of fronds, there is evolved a
structure such as the above-delineated species possess? Whether this
is the interpretation to be given of these plants, we shall best see
when considering whether it is the interpretation to be given of
plants which rank above them. Thus far we have dealt only with the
Cryptogamia. We have now to deal with the Phanerogamia or Phænogamia.
CHAPTER III.
THE MORPHOLOGICAL COMPOSITION OF PLANTS,
_CONTINUED_.
§ 187. That advanced composition arrived at in the _Archegoniatæ_, is
carried still further in the Flowering Plants. In these most-elevated
vegetal forms, aggregation of the third order is always distinctly
displayed; and aggregates of the fourth, fifth, sixth, &c., orders are
very common.
Our inquiry into the morphology of these flowering plants, may be
advantageously commenced by studying the development of simple leaves
into compound leaves. It is easy to trace the transition, as well as
the conditions under which it occurs; and tracing it will prepare us
for understanding how, and when, metamorphoses still greater in degree
take place.
§ 188. If we examine a branch of the common bramble, when in flower
or afterwards, we shall not unfrequently find a simple or undivided
leaf, at the insertion of one of the lateral flower-bearing axes,
composing the terminal cluster of flowers. Sometimes this leaf is
partially lobed; sometimes cleft into three small leaflets. Lower down
on the shoot, if it be a lateral one, occur larger leaves, composed
of three leaflets; and in some of these, two of the leaflets may be
lobed more or less deeply. On the main stem the leaves, usually still
larger, will be found to have five leaflets. Supposing the plant to be
a well-grown one, it will furnish all gradations between the simple,
very small leaf, and the large composite leaf, containing sometimes
even seven leaflets. Figs. 50 to 64, represent leading stages of the
transition. What determines this transition? Observation shows that
the quintuple leaves occur where the materials for growth are supplied
in greatest abundance; that the leaves become less and less compound,
in proportion to their remoteness from the main currents of sap; and
that where an entire absence of divisions or lobes is observed, it is
on leaves within the flower-bunch: at the place, that is, where the
forces which cause growth are nearly equilibrated by the forces which
oppose growth; and where, as a consequence, gamogenesis is about to
be set in (§ 78). Additional evidence that the degree of nutrition
determines the degree of composition of the leaf, is furnished by
the relative sizes of the leaves. Not only, on the average, is the
quintuple leaf much larger in its total area than the triple leaf;
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