The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
these foliaceous shoots, one on another; and where, as a consequence,
the first-formed of them become the main stem that acts as support
to secondary and tertiary stems; they lose their green, succulent
character, acquire bark, and become woody. In resuming the functions
of axes they resume the structures of axes, from which they had
deviated. In Fig. 71 are shown some of the leaf-like axes of _Rhipsalis
rhombea_ in their young state; while Fig. 72 represents the oldest
portion of the same plant, in which the foliaceous characters are
quite obliterated, and there has resulted an ordinary stem-structure.
One further fact is to be noted. At the same time that their
leaf-like appearances are lost, the axes also lose their separate
individualities. As they become stem-like, they also become integrated;
and they do this so effectually that their original points of junction,
at first so strongly marked, are effaced, and a consolidated trunk is
produced.
[Illustration: Figs. 71–72.]
Joined with the facts previously specified, these facts help us to
conceive how, in the evolution of flowering plants in general, the
morphological components that were once distinct, may become extremely
disguised. We may rationally expect that during so long a course of
modification, much greater changes of form, and much more decided
fusions of parts, have taken place. Seeing how, in an individual
plant, the single leaves pass into compound leaves, by the development
of their veins into mid-ribs while their petioles begin to simulate
axes; and seeing that leaves ordinarily exhibiting definitely-limited
developments, occasionally produce other leaves from their edges; we
are led to suspect the possibility of still greater changes in foliar
organs. When, further, we find that within the limits of one natural
order, petioles usurp the functions and appearances of leaves, at
the same time that in other orders, as in _Ruscus_, lateral axes so
simulate leaves that their axial nature would by most not be suspected,
did they not bear flowers on their mid-ribs or edges; and when, among
Cactuses, we perceive that such metamorphoses and re-metamorphoses
take place with great facility; our suspicion that the morphological
elements of Phænogams admit of profound transformations, is deepened.
And then, on discovering how frequent are the monstrosities which do
not seem satisfactorily explicable without admitting the development
of foliar organs into axial organs; we become ready to entertain
the hypothesis that during the evolution of the phænogamic type, the
distinction between leaves and axes has arisen by degrees.
With our preconceptions loosened by such facts, and carrying with us
the general idea which such facts suggest, let us now consider in what
way the typical structure of a flowering plant may be interpreted.
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