The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 192. To proceed methodically, we must seek a clue to the structures
of Phanerogams, in the structures of those inferior plants that
approach to them--_Archegoniatæ_. The various divisions of this
class present, along with sundry characters which ally them with
Thallophytes, other characters by which the phænogamic structure is
shadowed forth. While some of the inferior _Hepaticæ_ or Liverworts,
severally consist of little more than a thallus-like frond, among
the higher members of this group, and still more among the Mosses
and Ferns, we find a distinctly marked stem.[8] Some Archegoniates
(or rather Rhizoids) have foliar expansions that are indefinite in
their forms; and some have quite definitely-shaped leaves. Roots are
possessed by all the more-developed genera of the class; but there are
other genera, as _Sphagnum_, which have no roots. Here the fronds are
formed of only a single layer of cells; and there a double layer gives
them a higher character--a difference exhibited between closely-allied
genera of one group, the Mosses. Equally varied are the developments
of the foliar organs in their detailed structures: now being without
mid-ribs or veins; now having mid-ribs but no veins; now having
both mid-ribs and veins. Nor must we omit the similarly-significant
circumstance, that whereas in the lower Archegoniates the reproductive
elements are immersed here and there in the thallus-like frond, they
are, in the higher orders, seated in well-specialized and quite
distinct fructifying organs, having analogies with the flowers of
Phænogams. Thus, many facts imply that if the Phænogamic type is to
be analyzed at all, we must look among the Archegoniates for its
morphological components, and the manner of their integration.
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