The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
from the plant, and grow into new plants, as in _Mnium androgynum_
among the Mosses.” Now in the way above explained, these propagative
cells and proliferous buds, may continue developing in connexion with
the parent to various degrees before separating; or the buds which
are about to become fructifying organs may similarly, under increased
nutrition, develop into young fronds. As Sir W. Hooker says of the male
fructification in _Metzgeria furcata_,--“It has the appearance of being
a young shoot or innovation (for in colour and texture I can perceive
no difference) rolled up into a spherical figure.” On finding in this
same plant, that sometimes the proliferously-produced frond buds out
from itself another frond before separating from the parent, as shown
in Fig. 46, it becomes clear that this long-continued connexion may
readily pass into permanent connexion. And when we see how, even among
Phænogams, buds may either detach themselves as bulbils, or remain
attached and become shoots; we can scarcely doubt that among inferior
plants, less definite in their modes of organization, such transitions
must continually occur.
[Illustration: Figs. 73–76.]
Let us suppose, then, that Fig. 73 is the frond of some primitive
Archegoniate, similar in general characters to _Pellia epiphylla_,
Fig. 43; bearing, like it, the fructifying buds on its upper surface,
and having a slightly-marked mid-rib and rootlets. And suppose
that, as shown, a secondary frond is proliferously produced from
the mid-rib, and continues attached to it. Evidently the ordinary
discontinuous development, can thus become a continuous development,
only on condition that there is an adequate supply, to the secondary
frond, of such materials as are furnished by the rootlets: the
remaining materials being obtainable by itself from the air. Hence,
that portion of the mid-rib lying between the secondary frond and
the chief rootlets, having its function increased, will increase in
bulk. An additional consequence will be a greater concentration of
the rootlets--there will be extra growth of those which are most
serviceably placed. Observe, next, that the structure so arising
is likely to be maintained. Such a variation implying, as it does,
circumstances especially favourable to the growth of the plant, will
give to the plant extra chances of leaving descendants; since the area
of frond supported by a given area of the soil, being greater than
in other individuals, there may be a greater production of spores.
And then, among the more numerous descendants thus secured by it, the
variation will give advantages to those in which it recurs. Such a
mode of growth having, in this manner, become established, let us ask
what is next likely to result. If it becomes the habit of the primary
frond to bear a secondary frond from its mid-rib, this secondary frond,
composed of physiological units of the same kind, will inherit the
habit; and supposing that the supply of mineral matters obtained by
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