The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Already we have seen among the lower _Cryptogamia_, how, as they became
integrated and definitely limited, aggregates acquire the habit of
budding out other aggregates, on reaching certain stages of growth.
Cells produce other cells endogenously or exogenously; and fronds
give origin to other fronds from their edges or surfaces. We have
seen, too, that the new aggregates so produced, whether of the first
order or the second order, may either separate or remain connected.
Fissiparously-multiplying cells in some cases part company, while in
other cases they unite into threads or laminæ or masses; and fronds
originating proliferously from other fronds, sometimes when mature
disconnect themselves from their parents, and sometimes continue
attached to them. Whether they do or do not part, is clearly determined
by their nutrition. If the conditions are such that they can severally
thrive better by separating after a certain development is reached,
it will become their habit then to separate; since natural selection
will favour the propagation of those which separate most nearly at
that time. If, conversely, it profits the species for the cells or
fronds to continue longer attached, which it can only do if their
growths and subsequent powers of multiplication are thereby increased,
it must happen, through the continual survival of the fittest, that
longer attachment will become an established characteristic; and,
by persistence in this process, permanent attachment will result
when permanent attachment is advantageous. That disunion is really
a consequence of relative innutrition, and union a consequence of
relative nutrition, is clear _à posteriori_. On the one hand, the
separation of the new individuals, whether in germs or as developed
aggregates, is a dissolving away of the connecting substance; and
this implies that the connecting substance has ceased to perform its
function as a channel of nutriment. On the other hand, where, as
we see among Phænogams, there is about to take place a separation
of new individuals in the shape of germs, at the point where the
nutrition is the lowest, a sudden increase of nutrition will cause the
impending separation to be arrested; and the fructifying elements,
reverting towards the ordinary form, thereupon develop in connexion
with the parent. Turning to the Archegoniates, we find among them
many indications of this transition from discontinuous development
to continuous development. Thus the Liverworts give origin to new
plants by cells which they throw off from their surfaces; as, indeed,
we have seen that much higher plants do. “According to Bischoff,”
says Schleiden, “both the cells of the stem (_Jungermannia_ [now
_Lophocolea_] _bidentata_) and those of the leaves (_J. exsecta_)
separate themselves as propagative cells from the plant, and isolated
cells shoot out and develop while still connected with the parent
plant into small cellular bodies (_Metzgeria furcata_), which separate
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