For the sake of completeness it may be remarked, only by the way, that
the type of the proper harmonious-equipotential system may go hand in
hand with another type of “systems” which play a part in morphogenesis;
a type which we have shortly mentioned already and which will be studied
fully a few chapters later. We know that there are equipotential systems
with complex potencies: that is to say, systems which may produce a
whole organism equally well from any one of their elements; we know the
cambium of Phanerogams to be such a system. Now it is easily understood
that the germ of our Echinus, say in the stage of two or four or eight
cleavage cells, is not only an harmonious-equipotential system, but
a complex-equipotential system too. Not only may there arise a whole
organism out of 2/4 or 3/4 or 3/8, 4/8, 5/8, 6/8, 7/8 of its elements,
in which cases the harmonious rôle of the single element with regard to
its single performance in a totality is variable, but there may also
arise four whole single larvae out of the four cells of the four-cell
stage, or eight single whole larvae out of the eight-cell stage.[67]
In these cases, of course, each of the four or eight elements has
performed not a part of the totality, changing with its “position,” but
the totality itself. With respect to these possible performances the
“systems” present in the four or eight-cell stages of cleavage must be
called complex-equipotential ones.
[67] The eight larvae would be incomplete in some respect, but not with
regard to symmetry. They would be “whole” ones, only showing certain
defects in their organisation. See page 65 note 1, and page 73.
We propose to give the name of *mixed-equipotential systems* to all
those equipotential systems which, at the same time, may be regarded
as belonging to the harmonious or to the complex type. It is not only
among cleavage-stages that they are to be found; you may also find them
very clearly exhibited in our ascidian *Clavellina* for instance. We
know already that the branchial apparatus of this form is typically
harmonious-equipotential, but it is complex-equipotential too, for it
also may regenerate what is wanting in the proper way, by a budding
from the wound; and the same is true of many other cases, the flatworm
*Planaria* for instance.
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