You now, I hope, will have a general idea at least of the processes
of which the individual development of an animal consists. Of course
the specific features leading from the egg to the adult are different
in each specific case, and, in order to make this point as clear as
possible, I shall now add to our description a few words about what may
be called a comparative descriptive embryology.
COMPARATIVE EMBRYOLOGY
Even the cleavage may present rather different aspects. There may be a
compact blastula, not one surrounded by only one layer of cells as in
Echinus; or bilaterality may be established as early as the cleavage
stage--as in many worms and in ascidians--and not so late as in Echinus.
The formation of the germ layers may go on in a different order and
under very different conditions: a rather close relative of our Echinus,
for instance, the starfish, forms first the endoderm and afterwards
the mesenchyme. In many cases there is no tube of cells forming the
“endoderm,” but a flat layer of cells is the first foundation of all the
intestinal organs: so it is in all birds and in the cuttlefish. And,
as all of you know, of course, there are very many animal forms which
have no proper “larval” stage: there is one in the frog, the well-known
“tadpole,” but the birds and mammals have no larvae; that is to say,
there is no special stage in the ontogeny of these forms which leads an
independent life for a certain time, as if it were a species by itself,
but all the ontogenetical stages are properly “embryonic”--the germ is
always an “embryo” until it becomes the perfect young organism. And you
also know that not all skeletons consist of carbonate of calcium, but,
that there are skeletons of silicates, as in Radiolaria, and of horny
substance, as in many sponges. And, indeed, if we were to glance at the
development of plants also, the differences would seem to us probably
so great that all the similarities would seem to disappear.
But there are similarities, nevertheless, in all development, and we
shall now proceed to examine what they are. As a matter of fact, it was
especially for their sake that we studied the ontogeny of a special
form in such detail; one always sees generalities better if one knows
the specific features of at least one case. What then are the features
of most general and far-reaching importance, which may be abstracted
from the individual history of our sea-urchin, checked always by the
teachings of other ontogenies, including those of plants?
THE FIRST STEPS OF ANALYTICAL MORPHOGENESIS
Public-domain text, read in full here on John Shaqi.
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