If we look back upon the long fight of the schools of embryologists in
the eighteenth century about the question whether individual development
was to be regarded as a real production of visible manifoldness or as
a simple growth of visibly pre-existing manifoldness, whether it was
“epigenesis” or “evolutio,” there can be no doubt, if we rely on all
the investigations of the last hundred and fifty years, that, taken in
the descriptive sense, the theory of epigenesis is right. Descriptively
speaking there *is* a production of visible manifoldness in the course
of embryology: that is our first and main result. Any one possessed of
an average microscope may any day convince himself personally that it is
true.
In fact, true epigenesis, in the descriptive sense of the term, does
exist. One thing is formed “after” the other; there is not a mere
“unfolding” of what existed already, though in a smaller form; there is
no “evolutio” in the old meaning of the word.
The word “evolution” in English usually serves to denote the theory of
descent, that is of a real relationship of all organisms. Of course we
are not thinking here of this modern and specifically English meaning of
the Latin word *evolutio*. In its ancient sense it means to a certain
degree just the opposite; it says that there is no formation of anything
new, no transformation, but simply growth, and this is promoted not for
the race but for the individual. Keeping well in mind these historical
differences in the meaning of the word “evolutio,” no mistakes, it seems
to me, can occur from its use. We now shall try to obtain a few more
particular results from our descriptive study of morphogenesis, which
are nevertheless of a general bearing, being real characteristics of
organic individual development, and which, though not calculated of
themselves to further the problem, will in any case serve to prepare for
a more profound study of it.
The totality of the line of morphogenetic facts can easily be resolved
into a great number of distinct processes. We propose to call these
“elementary morphogenetic processes”; the turning in of the endoderm
and its division into three typical parts are examples of them. If we
give the name “elementary organs” to the distinct parts of every stage
of ontogeny which are uniform in themselves and are each the result of
one elementary process in our sense, we are entitled to say that each
embryological stage consists of a certain number of elementary organs.
The mesenchyme ring, the coelum, the middle-intestine, are instances of
such organs. It is important to notice well that the word elementary is
always understood here with regard to visible morphogenesis proper and
does not apply to what may be called elementary in the physiological
sense. An elementary process in our sense is a very distinct act of
form-building, and an elementary organ is the result of every one of
such acts.
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