The large and deep pit situated between the nostril and the eye (loreal
pit) in the Crotaline Viperidæ--whence the name Pit-vipers, or that of
"cuatro naricas" which is bestowed on them by the Spaniards of
Mexico--is divided into two chambers: an outer with large external
orifice, and an inner, rather more posterior in position and occupying
an excavation on the outer face of the maxillary bone. The inner walls
of these chambers are very thin and membranous, and form a partition
separating the two, except for the presence of a minute opening; this
partition is stretched across the hollow of the maxillary bone like the
membrane of a drum, and is supplied with blood-vessels and nerves, the
latter terminating in cells of variable form. The use of the organ, thus
situated at the base of the poison fang, and therefore in close
proximity to the sphincter of the poison duct, is still unknown.
Several of the Boidæ, such as _Python_ and _Corallus_, have deep pits in
some of the upper and lower labial shields, or also on each side of the
rostral shield; these problematic organs are in all probability also
sensory.
------------------------------------------------------------------------
CHAPTER VIII
VISCERA
In most snakes there is a very marked asymmetry of the viscera and their
blood-supply, the organs of the right side being anterior to, as well as
larger than, those of the left.
The heart in most cases is situated between the anterior seventh and the
anterior fourth of the body; it may be much farther back, beyond the
anterior third, in _Doliophis_, _Platurus_, and some Viperidæ and
Amblycephalidæ, in the middle in _Chersydrus_. It is of rather elongate
form, enclosed in a pericardium in which it lies freely, and has a sinus
venosus, two auricles, and a single ventricle divided by a septum. Three
arteries leave the ventricle, the pulmonary and two systemic arches. The
right systemic arch gives off the carotid artery, which in many snakes,
the common Grass-snake for instance, may branch into two, or in others
be double from its origin. The anterior abdominal vein is single in most
snakes, double in some Boidæ, and conveys blood from the ventral
body-wall to the liver. The caudal vein is continued as the renal
portal. Veins which have been regarded as remains of the two posterior
cardinal of lower Vertebrates have been found in some of the Boidæ.
The bifurcate transverse processes of the vertebræ at the limit between
the body and tail enclose the lymph-hearts, which are large and more or
less elongate, metamerically divided into several chambers, the right
often more developed than the left. The thymus gland lies on each side
of the trachea, near the heart, and the thyroid gland is in the middle
line, close to the base of the carotid artery.
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