The trachea is long, and the tracheal rings may be complete in front and
incomplete behind, or incomplete throughout. The bronchus opens at once
into the more or less elongate, usually single lung, with or without a
rudiment of a second, which seems to be constantly the left; in some
snakes the lung extends nearly to the cloacal region. In most of the
Boidæ there are two well-developed lungs, the left shorter than the
right. The lung has highly cellular walls in front, and becomes
thin-walled, smooth, or but little vascular, behind, where it may
receive its blood from the systemic and not from the pulmonary
circulation. In the Typhlopidæ and Viperidæ, as well as in some of the
Boidæ, Colubridæ, and Amblycephalidæ, the posterior end or the greater
part of the trachea may have its wall enlarged and provided with air
cells, resembling the normal lung, with which it is usually continuous;
this has been called the "tracheal lung," but, although serving as an
accessory breathing organ, it is not a prolongation of the true lung,
nor does it represent the missing left lung, as has been believed by
some authors.
The glottis has a longitudinal slit, and can be projected forwards when
the pharynx is obstructed by a voluminous prey. An epiglottis is usually
absent, or represented by a rudiment. It is, however, present in some
large American species of _Coluber_ (_Pityophis_), said to produce, when
hissing, a loud and hoarse sound which has been compared to the bellow
of the bull--hence the popular name of Bull-snakes by which they are
known. It has also been found in a few allied species from Mexico, for
which the genus _Epiglottophis_ has been proposed. This epiglottis is a
narrow, thin flap, erect in front of the glottis; it is not hinged, and
therefore not capable of falling down to cover the opening of the
windpipe during the process of swallowing, its function evidently being
to increase the sound produced by the escape of the air from the
windpipe.
The larynx is represented by two longitudinal bands of cartilage, united
by transverse bands; it is extremely long in some snakes
(_Leptognathus_).
The oesophagus, which may be extremely elongate, sometimes measuring
almost one-third of the digestive canal, passes into the tubular or
sac-like stomach, often with thickened walls, which itself gradually or
abruptly merges into the narrower intestine. The windings of the small
intestine are connected by ligamentous tissue, and enclosed in a common
sheath of peritoneum. In several of the Glyphodont Water-snakes
(Homalopsinæ and Hydrophiinæ), the intestine is much convoluted; in
_Herpeton_ it is even longer than the body, although when coiled
occupying only one-fourth of that length. The rectum is sometimes very
short, sometimes rather long, and its anterior portion may have a short
cæcum; it may be divided by transverse septa, with median or lateral
perforation.
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