Venoms: Venomous Animals and Antivenomous Serum-therapeuticsCalmette, A. (Albert)
Science
Venoms: Venomous Animals and Antivenomous Serum-therapeutics
Calmette, A. (Albert)
Antitoxins; Poisonous snakes; Toxins; Venom
With the exception of the species of _Atractaspis_, these snakes
are all ovoviviparous. The majority are terrestrial; a few lead a
semi-aquatic existence, while others are arboreal.
Their distribution includes Europe, Asia, Africa (with the exception
of Madagascar), and North and South America. They do not exist in
Australia.
They are divided into two _Sub-families_:--
A. The _Viperinæ_, in which the head, which is very broad and covered
with little plates and scales, has no pit between the nose and the eyes;
B. The _Crotalinæ_ (κρὁταλον, a rattle), in which the head is
incompletely covered with scales, and exhibits a deep pit on each side,
between the eye and the nostril.
* * * * *
Among snakes, the characters that serve as a basis for the
determination of genera and species are the general shape of the body,
especially that of the head, the arrangement of the cephalic scales,
the cranial skeleton, and the dentition.
* * * * *
_Cranial Skeleton._--The cranium is composed of a certain number of
bones, the homologues of which are found in the mammalian skeleton; but
the bones are complex, and subject to modifications according to the
structure and habitat of each species.
The special arrangement of the bones of the face is above all
characteristic of the poisonous snakes. Those forming the upper jaw,
the palate and the mandibles or “inter-maxillaries” are movable upon
each other and on the cranium. The upper and lower maxillaries are
united by an extensile ligament and articulated with the tympanic
bone, which permits the mouth to be opened very widely when the animal
swallows its prey.
* * * * *
_Dentition._--The _non-poisonous_ snakes have two rows of teeth in
the upper jaw--one external, the _maxillary_, usually composed of
from 35-40 small, backwardly curved teeth; the other internal, the
_palatine_, which only numbers from 20-22 teeth, having the same
curvature (fig. 1, A).
[Illustration: FIG. 1.--A, Cranial skeleton of one of
the non-poisonous _Colubridæ_ (_Ptyas mucosus_); B, cranial
skeleton of one of the poisonous _Colubridæ_ (_Naja tripudians_);
C, cranial skeleton of one of the poisonous _Colubridæ_
(_Bungarus fasciatus_); D, cranial skeleton of one of the
_Viperidæ_ (_Vipera russellii_); E, cranial skeleton of one
of the _Viperidæ Crotalinæ_ (_Crotalus durissus_); F, cranial
skeleton of one of the _Colubridæ Hydrophiinæ_ (_Hydrophis pelamis_).]
In the _poisonous_ snakes the maxillary bones are shorter, and the
_outer_ row is represented by a single long and tubular or grooved
tooth (the fang), fused with the maxillary bone, which is itself
movable (fig. 1, B, C, D, E,
F).
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