Venoms: Venomous Animals and Antivenomous Serum-therapeuticsCalmette, A. (Albert)
Science
Venoms: Venomous Animals and Antivenomous Serum-therapeutics
Calmette, A. (Albert)
Antitoxins; Poisonous snakes; Toxins; Venom
[Illustration: FIG. 2.--A, Maxillary bone and fangs
of one of the _Viperidæ_ (_Vipera russellii_); B, maxillary
bone and fangs of one of the _Colubridæ_ (_Naja tripudians_);
C, maxillary bone and fangs of one of the _Colubridæ_
(_Bungarus fasciatus_); D, maxillary bone and teeth of one
of the non-poisonous _Colubridæ_ (_Ptyas mucosus_). (After Sir Joseph
Fayrer.)]
Certain species (_Dipsas_) have maxillary teeth which increase in size
from front to rear; the longest teeth are _grooved_ and serve for the
better retention of prey, and also to impregnate it with saliva; but
they are not in communication with the poison-glands.
The _poison-fangs_ are normally covered with a fold or capsule of
mucous membrane, in which they are sheathed. This fold conceals a whole
series of _reserve_ teeth in different degrees of development, which
eventually become attached to the extremity of the maxillary when the
principal tooth falls out or is broken (fig. 2).
[Illustration: FIG. 3.--A, Fang of one of the
_Viperidæ_ (_Vipera russellii_); D, transverse section of the
fang.]
_Poison Apparatus._--The grooved or furrowed teeth in the
_Proteroglypha_ and the canaliculate teeth in the _Solenoglypha_ are
arranged, not for the purpose of seizing prey, but in order to deal it
a mortal blow by injecting the venom into its flesh.
In the normal position they lie almost horizontally, and exhibit no
mobility of their own. But, when the animal prepares to bite, their
erection is effected by the snake throwing its jaw back; and this
movement, which is always very sudden, enables it at the same time to
compress its poison-glands, by the aid of special constrictor muscles.
On examining the various species of poisonous snakes, we observe very
sharply marked differences in the arrangement and dimensions of the
teeth. Thus, in the VIPERIDÆ they are long, extraordinarily
sharp, and capable of producing deep wounds (fig. 2, A, and
fig. 3). They are traversed by an almost completely closed canal, from
the base, which communicates with the poison-duct, to the neighbourhood
of the point, where it opens very obliquely on the convex surface (fig.
3, A and D).
[Illustration: FIG. 4.--B, Fang of one of the
_Colubridæ_ (_Naja tripudians_); E, transverse section.]
[Illustration: FIG. 5.--C, Fang of one of the
_Hydrophiinæ_ (_Hydrophis pelamis_); F, transverse section.]
[Illustration: FIG. 6.--Three transverse sections of a
poison-fang of one of the _Colubridæ_, through A A, B
B, C C; P P, pulp cavity; V V V,
poison-canal (groove). (After C. J. Martin.)]
* * * * *
In the _Elapinæ_ sub-family of the _Colubridæ_, and especially in
the _Hydrophiinæ_, the teeth are much shorter and simply grooved or
canaliculate; that is to say, the canal communicates with the exterior
throughout its extent by a narrow slit, which traverses the entire
convex surface of the tooth (figs. 4, 5, and 6).
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