The growth of the wood of exogenous trees takes place thru the
ability, already noted, of protoplasmic cells to divide. The cambium
cells, which have very thin walls, are rectangular in shape, broader
tangentially than radially, and tapering above and below to a chisel
edge, Fig. 11. After they have grown somewhat radially, partition
walls form across them in the longitudinal, tangential direction,
so that in place of one initial cell, there are two daughter cells
radially disposed. Each of these small cells grows and re-divides, as
in Fig. 12. Finally the innermost cell ceases to divide, and uses its
protoplasm to become thick and hard wood. In like manner the outermost
cambium cell becomes bast, while the cells between them continue to
grow and divide, and so the process goes on. In nearly all stems,
there is much more abundant formation of wood than of bast cells. In
other words, more cambium cells turn to wood than to bast.
[Illustration: Fig. 11. Diagram Showing Grain of Spruce Highly
Magnified. PR, pith rays; BP, bordered pits; Sp W, spring wood; SW,
summer wood; CC, overlapping of chisel shaped ends.]
[Illustration: Fig. 12. Diagram Showing the Mode of Division of the
Cambium Cells. The cambium cell is shaded to distinguish it from the
cells derived from it. Note in the last division at the right that
the inner daughter cell becomes the cambium cell while the outer cell
develops into a bast cell. _From Curtis: Nature and Development of
Plants._]
In the spring when there is comparatively little light and heat, when
the roots and leaves are inactive and feeble, and when the bark, split
by winter, does not bind very tightly, the inner cambium cells produce
radially wide wood cells with relatively thin walls. These constitute
the spring wood. But in summer the jacket of bark binds tightly, there
is plenty of heat and light, and the leaves and roots are very active,
so that the cambium cells produce thicker walled cells, called summer
wood. During the winter the trees rest, and no development takes
place until spring, when the large thin-walled cells are formed again,
making a sharp contrast with those formed at the end of the previous
season.
It is only at the tips of the branches that the cambium cells grow
much in length; so that if a nail were driven into a tree twenty years
old at, say, four feet from the ground, it would still be four feet
from the ground one hundred years later.
Public-domain text, read in full here on John Shaqi.
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