The Foot-prints of the Creator : $b or, The Asterolepis of Stromness
John Stuart Mill · en
are in them rudimental. Their tails are finned on the under side only,—an
admitted feature of the salmon in an embryonic stage; and the mouth is
placed on the under side of the head,—also a mean and embryonic feature
of structure. These characters are essential and important, whatever
the Edinburgh Reviewer may say to the contrary; they are the characters
which, above all, I am chiefly concerned in looking to, for they are
features of embryonic progress, and embryonic progress is the grand key
to the theory of development.”
Such is the ingenious piece of special pleading which this most popular
of the Lamarckians directs against the standing and organization of the
earlier fishes. Let us examine it somewhat in detail, and see whether
the slight admixture of truth which it contains serves to do aught more
than to render current, like the gilding of a counterfeit guinea spread
over the base metal, the amount of error which lies beneath. I know
not a better example than that which it furnishes, of the entanglement
and perplexity which the meshes of an artificial classification, when
converted, in argumentative processes, into symbols and abstractions, are
sure to involve subjects simple enough in themselves.
Fishes, according to the classification of a preponderating majority
of the ichthyologists that have flourished from the earliest times
down to those of Agassiz, have been divided into two great series, the
_Ordinary_ or osseous, and the _Chondropterygii_ or cartilaginous. And
these two divisions of the class, instead of being ranged consecutively
in a continuous line, the one in advance of the other, have been
ranged in two parallel lines, the one directly abreast of the other.
There is this further peculiarity in the arrangement, that the line
of the cartilaginous series, from the circumstance that some of its
families rise higher and some sink lower in the scale than any of the
ordinary fishes, outflanks the array of the osseous series at both
ends. The front which it presents contains fewer genera and species
than that of the osseous division; but, like the front of an army drawn
out in single file, it extends along a greater length of ground. And
to this long-fronted series of the cartilaginous, or, according to
Cuvier, _chondropterygian_ fishes, the Placoid families of Agassiz
belong,—among the rest, the Placoids of the Silurian formations, Upper
and Lower. But though all the Placoids of this latter naturalist be
cartilaginous fishes, all cartilaginous fishes are not Placoids. The
_Sturionidæ_ are cartilaginous, and are, as such, ranked by Cuvier
among the _Chondropterygii_, whereas Agassiz places them in his Ganoid
order. Many of the extinct fishes, too, such as the _Acanthodei_,
_Dipteridæ_, _Cephalaspidæ_, were, as we have seen, cartilaginous in
their internal framework, and yet true Ganoids notwithstanding. The
principle of Agassiz’s classification wholly differs from that of Cuvier