A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
The alimentary canal is simple, but can be distinctly divided
into--(1st) an œsophagus, singular from the bell-shaped expansion
of its lower end; (2d) the stomach, which is directed forwards and
then doubled back; and (3d) the rectum. There is no distinct liver.
The circulation is lacunal. In one family there are well-developed
branchiæ, which differ entirely in their homologies and position
from these organs in all other Crustaceans. In the nervous system,
the sub-œsophageal ganglions vary in concentration from that degree
observed in the lower Macroura, to that in the highest Brachyoura; but
the supra-œsophageal ganglions are always much less concentrated, and
are even embryonic in condition; presenting a difference not observed
in other Crustaceans. On the under side of the sub-œsophageal ganglion,
two nerves, apparently splanchnic, arise, and run almost parallel and
under the collar surrounding the œsophagus; they are very remarkable
from their great size, and from forming a plexus together with a
large branch, arising on each side from the collar close behind the
supra-œsophageal ganglion,--a structure unlike anything observed in
other Crustaceans. The eyes, as already remarked, are rudimentary, and
singular from being imbedded at a distance from the anterior end of the
animal. In the basal confluent segments of the outer maxillæ there are
two orifices, leading into little sacks, which I believe are olfactory
organs: again there are two other orifices on each side of the thorax,
beneath the first pair of cirri, leading into sacks, with a curious
elastic vesicle suspended within them; and these I can hardly doubt are
acoustic organs. Of these orifices and organs, there is no trace in the
same relative positions in any known Crustacean.
Cirripedes are ordinarily bisexual, in which they differ from all
Crustaceans: when the sexes are separate, the males are minute,
rudimentary in structure, and permanently epizoic on the females; to
these latter facts we have a partial analogy in some of the Suctorial
Entomostracans; but a far closer analogy in certain Rotifers, which are
considered by many naturalists as Crustaceans; but to the above subject
I shall almost immediately have to recur.
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