A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc. — John Shaqi
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
The male excretory organ is probosciformed and capable of the most
varied movements; it is single and medial; it is seated (in the one
instance in which this point can be safely judged of) at the extremity
of the abdomen, and therefore near the normal position of the anus;
in all these respects there is a very great difference from other
Crustaceans, in which the male organs are laterally double, and are not
seated at the extremity of the abdomen. In regard to the female organs,
the ovarian tubes and cæca inosculate together: there are no oviducts;
the ova, connected together by membrane, and so forming the "ovigerous
lamellæ," become exposed by the exuviation of the lining tunic of the
carapace or sack, and by the formation of a new tunic on the under side
of these lamellæ; a process, I believe, unknown in other Crustaceans.
The metamorphoses are highly complex. The larva in its first stage
bears a very close general resemblance, in having three pairs of
natatory appendages, the first being uniramous and the two others
biramous, and in having a single eye on its broad anterior front, to
the larvæ of most Entomostracans; but I cannot avoid the belief, that
this resemblance is only apparent, and not essential; and of false
resemblances, how many instances occur in the animal kingdom! In the
larva, when first freed from the egg, both pairs of antennæ are in
process of formation within envelopes: the mouth is probosciformed
and capable of movement, but is destitute of gnathites; it occupies a
position between the three pairs of natatory limbs; and these limbs I
must believe, for reasons hereafter to be assigned, answer (improbable
as I am well aware it must at first appear) to the second, third,
and fourth thoracic legs of the archetype Crustacean: the two hinder
pairs of limbs apparently soon become captorial, or fitted to secure
prey. Now, I cannot find in the published accounts of the larvæ of
Entomostracans, any that answer to this description.
The larva in the last stage might be included in the vast class of
Entomostracans: the attachment of the eyes to the singular apodemes
produced inwards from the basal segment of the great prehensile
antennæ, and the development of only the posterior six pairs of
thoracic limbs, are its chief peculiarities: but its rudimentary mouth,
owing to its transitional or pupal condition, renders the assignment of
its proper rank difficult.
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