A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
Having now given this short comparative sketch of the structure of
a Cirripede, I may venture to express strongly my opinion, that the
group is formed on a distinct type; as different from the other
three or four main Crustacean groups, namely, the Podophthalmia,
Edriophthalmia, Branchiopoda, and Entomostraca, as these differ from
each other; the differences, moreover, being of the kind considered
by the highest authorities on this subject, as the most important. It
should be observed that there is no special blending at either end of
the Cirripedial series, towards any one of the other main groups of
Crustacea; it is hardly possible to take some one Cirripede, and say
that it leads, more plainly than some other Cirripede, into ordinary
Crustaceans. Moreover, a great range of structure, as we shall soon
briefly show, is included within the group: I can adduce three or four
undoubted Cirripedes, very considerably more different from each other,
than any two members within the sub-class Podophthalmia, or within the
Edriophthalmia, or the Branchiopoda, and quite as different as within
the Entomostraca.
The opinion here expressed, that Cirripedes form a sub-class of equal
value with the other main Crustacean groups, I am well pleased to find,
accords with Mr. Dana's[13] view, who remarks that this sub-class
"has so many peculiarities of structure, that it should be regarded
as a distinct type, rather than a subordinate division of the third
(or Entomostracan) type." M. Milne Edwards,[14] after dividing all
Crustacea into two groups, divides one of them into four legions;
and of one of these, the Entomostraca, he makes the Cirripedes a
sub-group. I feel so entire a deference for any opinion on affinities
or classification expressed by Milne Edwards, that I differ from
him with the greatest hesitation. He does not give his reasons for
assigning so subordinate a rank to Cirripedes, but I imagine it is from
the nature of their metamorphoses: but if this be the case, I cannot
understand why he should assign to his Branchiopods a rank equal to his
Entomostracans. Moreover, I must repeat, that I do not believe that
the larvæ do resemble the larvæ of Entomostracans and Branchiopods
nearly so closely as at first appears to be the case. I may add, that
Burmeister[15] has assigned to the Cirripedes a place amongst the
Crustacea, almost equally subordinate to that given to them by Milne
Edwards.
[13] 'Crustacea: United States Exploring Expedition,' p. 1407, 1852.
[14] 'Annales des Sciences Nat.,' tom. xviii, p. 120, 1852.
[15] 'Beiträge zur Naturgesehichte der Rankenfüsser,' 1834.
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