A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc. — John Shaqi
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
That Cirripedes have some special affinity to the Entomostraca, may
be inferred from the fewness of the cephalic appendages, the biramous
legs, the state of the abdomen, and the form of the carapace. Perhaps
in the peculiar state of confluence of the lower segments of the
gnathites, in the aborted antennæ, the rudimentary eyes, and in the
minute parasitic males (when such exist), there is a more direct
relation to the Suctorial division of the Entomostraca; but some of
these resemblances are probably only analogical, resulting from the
fixed condition of both groups. It should not be overlooked, that
out of the three orders into which Cirripedes may be divided, in
the two latter, the mature animal presents hardly any resemblance
to an Entomostracan. From the distinct presence in either pupa or
mature animal of the fourteen segments of the cephalo-thorax; from
the apparent composition of the carapace, as will be subsequently
explained; and from the concentrated condition of the nervous system,
one is led to glance at the higher Crustacea; and here we shall find
amongst the Podophthalmia, one aberrant group of low organisation,
namely, that including Phyllosoma, Amphion, &c., in which more points
of resemblance to Cirripedes may be detected, than, as I believe, in
any other group whatever; for we here see that remarkable elongation
of the head in front of the mouth, so eminently characteristic of
Cirripedes; we have a carapace overlapping the thorax, which is
sometimes free beneath; we have the abdomen sometimes almost obsolete;
we have biramous legs: and especially we have the posterior cephalic
and the first thoracic appendages more or less rudimentary and
obsolete; and this, I infer from Mr. Dana, is a very rare phenomenon,
though characteristic of all _ordinary_ Cirripedes, in which the
seventh and eighth segments with their appendages have disappeared.
In the order including Phyllosoma, &c., namely, in the Macroura, the
ganglions which give nerves to the five posterior thoracic limbs, are
distinct from the great sub-œsophageal ganglion which supplies the
several anterior appendages; this is the case with those Cirripedes in
which all the infra-œsophageal ganglions are not concentrated into one.
In the Macroura and Brachyoura, the first pair of legs almost always
differs in structure from the others, so does the homologous or second
cirrus in Cirripedes differ from the four succeeding pairs; in some
few Macroura, the second leg is antenniformed, so in some few cases is
the homologous (or third) cirrus; J. Vaughan Thompson was even struck
by the resemblance in the curious, doubly pectinated spines on the
anterior limbs of Mysis (allied to Phyllosoma[16]), and on those of
many Cirripedes: these several latter resemblances may be small, but
certainly I do not believe that they are accidental. Now the little
group of Crustaceans, which includes Phyllosoma, &c., has lately
been placed, by Milne Edwards, as a satellite amongst the Macrourous
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