Fossil plants, Vol. 2 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 2 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In several species, grouped round the widely distributed type
_Lycopodium Selago_ Linn., the comparatively short, erect or suberect,
shoots form fairly compact tufts; the ordinary foliage-leaves function
as sporophylls, and the sporangia are not localised on special portions
of shoots. From this type, we pass to others in which the fertile
leaves tend to be confined to the tips of branches, but hardly differ
in form from the sterile. A further degree of specialisation is
exhibited by species with well-defined cones composed of leaves (or
bracts), the primary function of which is to bear sporangia and to
afford a protective covering to the strobilus[93].
_Lycopodium rufescens_ Hook. An Andian species with stout dichotomously
branched erect stems bears on the younger shoots crowded leaves with
their thick and broadly triangular laminae pointing upwards, but on the
older and thick shoots the laminae are strongly reflexed (fig. 121,
A). The lower part of the specimen represented in fig. 121, A, shows
tangentially elongated scars and persistent leaf-bases or cushions left
on the stem after the removal of the free portions of the leathery
leaves, a surface-feature which also characterises the Palaeozoic genus
_Lepidodendron_. The reflexed leaves and persistent leaf-cushions are
clearly seen in the piece of old stem of _Lycopodium dichotomum_ Jacq.,
a tropical American species reproduced in fig. 121, B. Such species
as _L. erythraeum_ Spring, and others with stiff lanceolate leaves
exhibit a striking resemblance to the more slender shoots of some
recent conifers, more especially _Araucaria excelsa_, _A. Balansae_,
_Cryptomeria_, _Dacrydium_ and other genera.
[Illustration: FIG. 121. _Lycopodium._
A. _Lycopodium rufescens._
B. _L. dichotomum._
C. _L. tetragonum._
D. _L. nummularifolium._
E. _L. Dalhousianum._
F. _L. casuarinoides._
G. _L. volubile._
(From specimens in the Cambridge Herbarium and Botanic Garden. M.S.)]
[Illustration: FIG. 122. _Lycopodium squarrosum._ The branches of the
larger shoot terminate in cones. (From a plant in the Cambridge
Botanic Garden. Reduced.)]
In _Lycopodium tetragonum_ Hook., (fig. 121, C), a species from the
Alpine region of the Andes, the long, pendulous and repeatedly forked
branches bear four rows of fleshy ovate leaves and simulate the
vegetative characters of certain conifers.
[Sidenote: LYCOPODIUM]
[Illustration: FIG. 123. _Lycopodium cernuum._
(From a specimen in the Cambridge Herbarium. ½ nat. size.)]
_L. squarrosum_ Forst. (fig. 122) a tropical species from India,
Polynesia, and other regions, is characterised by its stout stems
reaching a diameter of 2·5 cm., bearing long pendulous branches with
large terminal cones composed of sporophylls differing but slightly
from the foliage leaves. The plant represented in the photograph serves
as a good illustration of the practical identity in habit between
Palaeozoic and recent genera.
[Illustration: FIG. 124. _Lycopodium obscurum._]
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