On Germinal Selection as a Source of Definite Variation — John Shaqi
On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
Such are the objections raised against the theory of selection by its
opponents, and put forward as insurmountable obstacles to the process. Nor
are such objections relevant only in the case of protective colorings; they
are applicable in all cases where the process of selection is concerned.
Take the case of instincts that are called into action only once in life,
as, for example, the pupal performances of insects, the artificial
fabrication of cocoons, etc. How is it that the useful variations were
always present here? And yet they must have been present, if such
complicated spinning instincts could have taken their rise as are
observable in the silk-worm, or in the emperor-moth. And they have been
developed, and that in whole families, in forms varying in all species, and
in every case adapted to the special wants of the species.
Particularly striking is the proof afforded of this constant presence of
the useful variations by cases where we meet with the development of highly
special adaptations that are uncommon even for the group of organisms
concerned. Such a case, for example, is the apparatus designed for the
capture of small animals and their digestion, found in widely different
plants and widely separated families. On the other hand, very common
adaptations, such as the eyes of animals, show distinctly that in all cases
where it was necessary, the useful variations for the formation of {28} an
eye were presented, and were presented further exactly at spots at which
organs of vision could perform their best work: thus, in Turbellaria and
many other worms that live in the light, at the anterior extremity of the
body and on the dorsal surface; in certain mussels, on the edge of the
mantle; in terrestrial snails, on the antennae; in certain tropical marine
snails inhabiting shallow waters, on the back; and in the chitons even on
the dorsal surface of the shell!
But even taking the very simplest cases of selection, it is impossible to
do without this assumption, that the useful variations are always present,
or that _they always exist in a sufficiently large number of individuals
for the selective process_. You know the thickness and power of resistance
of the egg-shells of round-worms. The eggs of the round-worms of horses
have been known to continue their course of development undisturbed even
after they had been thrown into strong alcohol and all other kinds of
injurious liquids--much to the vexation of the embryologists, who wished to
preserve a definite stage of development and sought to kill the embryo at
that stage. Indeed, think of the result, if in the course of their
phylogenesis stout and resistant variations of egg-shells had not been
presented in these worms, or had not always been presented, or had not been
presented in every generation and not in sufficient quantities.
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