On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
have accidentally produced this brick-red _in a single case_, but that it
should have done so three times and in three species, which live together
but are otherwise not related, is a far more violent and improbable
assumption than that of a causal connexion of this coincidence. Now
hundreds of cases of such mimicry exist in which the color-tints of the
copy are met with again in more or less precise and sometimes in
exceedingly exact imitations, and there are thousands of cases in which the
color-tint of a bark, of a definite leaf, of a definite blossom, is
repeated _exactly_ in the protectively colored insect. In such cases there
can be no question of accident, but _the variations presented to personal
selection must themselves have been produced by the principle of the
survival of the_ {53} _fit!_ And this is effected, as I am inclined to
believe, through such profound processes of selection in the interior of
the germ-plasm as I have endeavored to sketch to you to-day under the title
of germinal selection.
I am perfectly well aware how schematic my presentation of this process is,
and must be at present, owing mainly to our inability to gain exact
knowledge concerning the fundamental germinal constituents here assumed.
But I regard its existence as assured, although I by no means underrate the
fact that eminent thinkers, like Herbert Spencer, contest its validity and
believe they are warranted in assuming a germ which is composed of _similar
units_. I strongly doubt whether even so much as a _formal_ explanation of
the phenomena can be arrived at in this manner. So far as direct
observation is concerned, the two theories stand on an equal footing, for
neither my dissimilar, nor Spencer's similar, units of germinal substance
can be _seen_ directly.
The attempt has been recently made to discredit my _Anlagen_, or
constitutional germ-elements, on the ground that they are simply a
subtilised reproduction of Bonnet's old theory of preformation.[18] This
{54} impression is very likely based upon ignorance of the real character
of Bonnet's theory. I will not go into further details here, particularly
as Whitman, in several excellently written and finely conceived essays, has
recently afforded opportunity for every one to inform himself on the
subject. My determinants and groups of determinants have nothing to do with
the preformations of Bonnet; in a sense they are the exact opposites of
them; they are simply _those living parts of the germ whose presence
determines the appearance of a definite organ of a definite character in
{55} the course of normal evolution_. In this form they appear to me to be
an absolutely necessary and unavoidable inference from the facts. There
_must_ be contained in the germ parts that correspond to definite parts of
the complete organism, that is, parts that constitute the reason why such
other parts are formed.
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