On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
when about to seize it, and is thus one of the causes which has led to
the wide extension and maintained the permanence of this now dominant
group. Those who believe that such peculiar structures can only have
arisen by very minute successive variations, each one advantageous to
its possessor, must see, in the possession of such an organ by one
group, and its complete absence in every other, a proof of a very
ancient origin and of very long-continued modification. And such a
positive structural addition to the organization of the family,
subserving an important function, seems to me alone sufficient to
warrant us in considering the Papilionidæ as the most highly developed
portion of the whole order, and thus in retaining it in the position
which the size, strength, beauty, and general structure of the perfect
insects have been generally thought to deserve.
Footnote 2:
Transactions of the Linnean Society, vol. xxiii. p. 495.
The Papilionidæ are pretty widely distributed over the earth, but are
especially abundant in the tropics, where they attain their maximum of
size and beauty and the greatest variety of form and colouring. South
America, North India, and the Malay Islands are the regions where these
fine insects occur in the greatest profusion, and where they actually
become a not unimportant feature in the scenery. In the Malay Islands in
particular the giant Ornithopteræ may be frequently seen about the
borders of the cultivated and forest districts, their large size,
stately flight, and gorgeous colouring rendering them even more
conspicuous than the generality of birds. In the shady suburbs of the
town of Malacca two large and handsome Papilios (_Memnon_ and
_Nephelus_) are not uncommon, flapping with irregular flight along the
roadway, or, in the early morning, expanding their wings to the
invigorating rays of the sun. In Amboyna and other towns of the
Moluccas, the magnificent _Deiphobus_ and _Severus_, and occasionally
even the azure-winged _Ulysses_, frequent similar situations, fluttering
about the orange-trees and flower-beds, or sometimes even straying into
the narrow bazaars or covered markets of the city. In Java the
golden-dusted _Arjuna_ may often be seen at damp places on the roadside
in the mountain districts, in company with _Sarpedon_, _Bathycles_, and
_Agamemnon_, and less frequently the beautiful swallow-tailed
_Antiphates_. In the more luxuriant parts of these islands one can
hardly take a morning’s walk in the neighbourhood of a town or village
without seeing three or four species of _Papilio_, and often twice that
number. No less than 120 species of the family are now known to inhabit
the Archipelago, and of these ninety-six were collected by myself.
Twenty-nine species are found in Borneo, being the largest number in any
one island, twenty-three species having been obtained by myself in the
vicinity of Sarawak; Java has twenty-seven species; Celebes and the
Peninsula of Malacca twenty-three each.
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