Cutting the frenula caused the pedalia to contract but seemed not to affect the ability to swim. Comparing the velarium of the Cubomedusæ with the velum of the Hydromedusæ, I recall no observations similar to the ones here noted, though it seems that the two may have quite similar functions. It seems somewhat probable that the velum, and also the velarium, may function in obtaining food,--and this besides their function in swimming. Their probable function in swimming, as is well known, is evidently to narrow the mouth of the bell and thus to cause the water to be forced out in a smaller but more rapid stream, giving the animal a steady and more prolonged movement through the water at every contraction of the bell. In regard to taking food, I observed that a small crustacean, in the process of being swallowed by an Olindiad, seemed to be held by the velum being firmly contracted about it while the proboscis was working itself over the crustacean. It would seem, furthermore, that my supposition is supported for Charybdea by the fact that the pedalia and tentacles were contracted so as to be brought inside the bell when the velarium was cut. The stimulus of cutting the velarium may be comparable to a stimulus from some object touching it, and thus cause the pedalia and tentacles to come reflexly to aid in capturing or holding the object, a fish, crustacean, or such, to be captured. _Pedalia, Interradial Ganglia, Tentacles_--Experiments 15, 23, 27-31, 41b.--When the pedalia were removed, the power of the animal to guide itself was completely gone. When one pedalium was cut the others contracted, while stroking the outer edge of the pedalia, touching the sensory clubs, or sharply pricking the subumbrella, often produced the same result. (See also Nerve.) The upper part of the subumbrella seemed not so sensitive and more seldom produced the reflex of the pedalia, while the base of the stomach did not give it at all. Stroking the outer edge of the pedalia of _Tripedalia cystophora_, the second of the two species of Cubomedusæ described by Conant, also caused the pedalia to be contracted inwards. I may note here that the muscle fibers under the ectoderm of the pedalia are specially well developed at and near the inner and outer edges, both in Charybdea and Tripedalia. On the flattened sides of the pedalia the muscle fibers are fewer. When the pedalia were cut off far enough up to remove the interradial ganglia, coördination was not affected and the animal could pulsate well enough but with little progress. (See above under Velarium and Frenula.) An isolated tentacle is capable of squirming contractions, and when stimulated at either end, it would contract wholly or in part only. The pedalia, then, it would seem, serve also as a steering apparatus, for which they are admirably fitted, considering their blade-like thinness.