Considering, now, the reflexes noted under this head and the preceding one, we find that there is an intimate nervous connection between the velarium and frenula, subumbrella, sensory clubs, nerve, and a single pedalium, on the one hand, and the pedalia on the other hand. This is born out fully, furthermore, by the histological evidence--(See Introduction and “Cubomedusæ”). Considering the subumbral plexus of ganglion cells and fibers, including the velarium and the frenula, which is in connection with the nerve ring and this again with the sensory clubs and the interradial ganglia at the bases of the pedalia, we have a basis for these reflexes. While Conant failed to demonstrate nerves (“Cubomedusæ”) from the interradial ganglia to the pedalia, yet, that a nervous connection exists between the pedalia and the bell is well shown by his physiological experiments. I have, furthermore, demonstrated ganglion cells under the ectoderm of the tentacles (see Histology). Romanes obtained quite similar results in the Hydromedusæ. He found that when a tentacle of Sarsia was slightly stimulated, it alone would contract, but when it was more strongly stimulated the other tentacles also would respond as also the manubrium. I find no evidence in Conant’s notes of any such response of the manubrium of Charybdea, except when the clubs were cut off. The reflex obtained on stimulating the subumbrella of Charybdea, when the pedalia would contract, is somewhat different from that obtained by Romanes, who found that the most sensitive part of the subumbrella in producing a reflex of the margin was at the junction of the manubrium to the bell and that the subumbrella below this point did not give the reflex. _Stomach, Suspensoria, Proboscis, Subumbrella_--Experiments 12, 18, 19, 24-26, 29, 31.--The proboscis and the stomach with the phacelli when cut out, contracted with or without the lips removed. The isolated lips also contracted (twitched). Pieces of the sides connected only with the stomach and suspensoria, or with the margin (Experiment 47 (?)) twitched spontaneously, but seldom did so when these were removed. In one instance the whole side was cut out so as to exclude the radial ganglion but still connected with a portion of the suspensorium. This pulsated, or contracted, but on being halved transversely, the lower half ceased to contract while the upper half connected with the suspensorium, continued to contract. Cutting off the whole stomach end of the animal excited to very rapid pulsations of the remaining part, with the stream of water stronger out the aboral end than past the velarium.