Professor Todd found from the examination of a small series of material that about an equal number of right-and left-handed flowers is produced. He also says: “It is also a fact of observation that the flowers of a cluster on any one branch and opening about the same time are either all right-handed or all left-handed. Any plant, however, if it is at all large, exhibits right-and left-handed flowers in about equal numbers.” The regularity with which the flowers are divided into the two classes is very striking. Table B shows the condition of ten plants observed at the same time. TABLE B. Plant I 7 pistils right-handed, 7 left-handed. " II 6 " " 6 " " III 8 " " 9 " " IV 29 " " 31 " " V 11 " " 7 " " VI 10 " " 7 " " VII 10 " " 13 " " VIII 3 " " 3 " " IX 3 " " 2 " " X 6 " " 9 " —— —— —— Total 10 93 pistils right-handed, 94 left-handed. So in these ten plants the number of right-and left-handed flowers is practically equal. The greatest difference in the number of the two kinds is seen in number X, where forty per cent. are right-handed and sixty per cent. left-handed. Considerable care was exercised in determining the number of right-and left-handed flowers opening on the racemes of different branches at the same time. Only those flowers were considered which had opened simultaneously. In order to effect this, all the flowers were removed from the plant the evening before and note was made of the condition of those opening the next day. The following diagram shows the conditions of flowers opening on three plants on the morning of August 20, braces indicating the branches of the plant, and the straight lines the racemes; the numbers of right- and left-handed flowers being indicated under the raceme by _r_ and _l_. (See page 20.) From the table, it will be seen that there are on the first plant 8 left-and 11 right-handed flowers; on the second, 24 left-and 27 right-handed; on the third, 7 left-and 9 right-handed flowers. The numbers of right-and left-handed flowers occurring on the divisions a and b of the main branches, A and B, of the three plants, are as follows: [Illustration] I.-Aa, 2_l_, 1_r_; Ab, 1_l_, 3_r_; Ba, 4_l_, 4_r_; Bb, 1_l_, 3_r_. II.-Aa, 8_l_, 13_r_; Ab, 13_l_, 11_r_; Ba, 1_l_, Bb, 1_l_; B, 2_l_, 3_r_. III.-Aa, 4_l_, 2_r_; Ab, 2_l_, 1_r_; Ba, 3_r_; Bb, 1_r_; B, 2_r_, 1_l_.