With equal emphasis, I must deny that the enclosed space in the ring represents any plane of division in the chromatin thread; and that the insertion of the spindle fibers is at any place except at the center of what would be the typical rod-shaped chromosome were the ring straightened out. We encounter in de Sinéty’s interpretation of these rings the very error against which I was careful to caution elsewhere in this paper, _i. e._, _of regarding the points where the fibers are attached as the crossed ends of a simple segment_. This mistake de Sinéty has made, and has thereby vitiated all his conclusions concerning the structure of the tetrads. It is not necessary to repeat here the proof which I have brought forward in support of my views. No one, I am sure, will find difficulty in reducing the various forms of chromosomes found in the first spermatocytes to the type of a doubly split rod, in which one plane of division is parallel to the long axis and the other at right angles to it. The explanation offered by de Sinéty requires us to conceive a doubly split rod in which one separating space may vary indefinitely while the other is constant. There is here no common type, but an infinitely variable one, which differs with every modification of the interspace between the first pair of chromatids in each chromosome. As a constructive basis for the foundation of his theory of a double longitudinal division, de Sinéty uses particularly the chromosomes of _Œdipoda_ (_Hippiscus_) _miniata_, represented in figures 129 and 130, concerning which he says: “Survient le phénomène exceptionnellement important de la seconde division longitudinale; nous regardons comme un point capital dans notre travail d’en mettre l’existence hors de doute et pour cela nous désirons ne faire appel qu’à des images extrêmement claires. Nous considérons comme telles les fig. 129 et 130 rapprochées l’une de l’autre. “Il est de toute évidence que le chromosome _a_, fig. 130, n’est que le chromosome de même désignation, fig. 129, dont les deux anses jumelles se sont clivées. De même, le chromosome en forme de boucle, _c_, fig. 129, dont les deux branches représentent, comme nous l’avons fait remarquer, deux anses jumelles, se retrouve avec un clivage très évident en _d_, fig. 123. On pourrait faire les mêmes rapprochements entre _b_, fig. 105, et _a_, fig. 107; ici, le clivage est moins avancé, mais les granules sont nettement divisés.”