The Principles of Biology, Volume 2 (of 2) — John Shaqi
The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
In these cases, however, the tertiary aggregate, so
far as it is formed, is but very feebly integrated; and its integration
is but temporary. For not only do these younger fronds that bud out
from the mid-ribs of older fronds, develop rootlets of their own; but
as soon as they are well grown and adequately rooted, they dissolve
their connexions with the parent-fronds, and become quite independent.
From these transitional forms we pass, in the higher _Jungermanniaceæ_,
to forms composed of many fronds that are permanently united by a
continuous stem. A more-developed aggregate of the third order is
thus produced. But though, along with increased definiteness in
the secondary aggregates, there is here an integration of them so
extensive and so regular, that they are visibly subordinated to the
whole they form; yet the subordination is really very incomplete.
In some instances, as in _Radula complanata_, Fig. 47, the leaflets
develop roots from their under surfaces, just as the primitive frond
does; and in the majority of the group, as in _J. capitata_, Fig. 48,
roots are given off all along the connecting stem, at the spots where
the leaflets or frondlets join it: the result being that though the
connected frondlets form a physical whole, they do not form, in any
decided manner, a physiological whole; since successive portions of
the united series, carry on their functions independently of the rest.
Finally, the most developed members of the group, whether lineally
descended from the less developed or from an early type common to the
two, present us with tertiary aggregates which are physiologically as
well as physically integrated.[5] Not lying prone like the kinds thus
far described, but growing erect, the stem and attached leaflets become
dependent upon a single root or group of roots; and being so prevented
from carrying on their functions separately, are made members of a
compound individual: there arises a definitely-established aggregate of
the third degree of composition.
The facts as arranged in the above order are suggestive. Minute
aggregates, or cells, the grouping of which we traced in § 182, showed
us analogous phases of indefinite union, which appeared to lead the
way towards definite union. We see here among compound aggregates, as
we saw there among simple aggregates, the establishment of a specific
form, and a size that falls within moderate limits of variation. This
passage from less definite extension to more definite extension,
seems in the one case, as the other, to be accompanied by the result,
that growth exceeding a certain rate, ends in the formation of a new
aggregate, rather than an enlargement of the old. And on the higher
stage, as on the lower, this process, irregularly carried out in the
simpler types, produces in them unions that are but temporary; while in
the more-developed types, it proceeds in a systematic way, and ends in
the production of a permanent aggregate that is doubly compound.
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